I'll give it a stab. You want to create the illusion of teeth, not draw teeth. When you outline around the entire tooth, it appears that there is a space between the teeth. What you want to do is draw the shape of the "not teeth" (i.e., the gum line, the inside of the mouth, tongue, bottom lip, etc.,) and what is left is the teeth. You can further delineate these "teeth" by noting shadow, the curve of the muzzle, and hitting a (slight) highlight, but you don't "draw teeth".
The "not teeth" concept will come as no surprise to my students and demo watchers. I also paint a "not nose" and a "not mouth".
Otherwise, you've done a terrific job! (I don't even see much of a problem with the teeth....)
Peggy
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